The husband of former First Minister Nicola Sturgeon has been charged in connection with the embezzlement of funds from the Scottish National Party.
Peter Murrell, 59, was taken into custody at 9.13am on Thursday and was questioned by Police Scotland detectives.
He was previously arrested as a suspect on 5 April 2023 before being released without charge.
BBC Scotland understands Murrell has resigned his SNP membership.
He was charged at 6.35pm after further questioning by officers investigating the funding and finances of the party as part of Operation Branchform.
Murrell has since been released from police custody at Falkirk Police Station and returned to his home near Glasgow shortly after 8pm.
He resigned as the SNP's chief executive in March of last year after 22 years in the job.
An SNP spokesperson said: "While this development will come as a shock, the police investigation remains ongoing and it would, therefore, be inappropriate to make any comment."
Police Scotland launched a formal investigation into the SNP's finances in July 2021 after receiving complaints about how donations were used.
Questions had been raised about more than £660,000 (NZ$1.3 million) in donations given to the party for use in a fresh independence referendum campaign.
When Murrell was arrested last year, police searched the house he shares with Sturgeon near Glasgow and the SNP headquarters in Edinburgh. The couple married in 2010.
The house was sealed off with blue and white tape, while a tent was erected on the driveway. Items were brought from the house to the tent, where the BBC understands a vehicle was parked.
Police also seized a camper van from outside the Dunfermline home of Murrell's mother.
The Niesmann and Bischoff vehicle, which can retail for more than £100,000, was seized by police the same morning that Murrell became the first senior party figure to be arrested in the probe.
On 18 April 2023, SNP treasurer and MSP Colin Beattie was arrested and interviewed by police before being released pending further investigation. He later resigned as treasurer.
Sturgeon was arrested on 11 June when she voluntarily arranged with Police Scotland to be questioned as part of the investigation, a spokesperson said at the time.
She was released without charge seven hours later pending further inquiries.
Sturgeon had unexpectedly announced she was resigning as SNP leader and first minister four months earlier in February 2023.
A spokesperson for the Crown Office and Procurator Fiscal Service (COPFS) said: "Senior professional prosecutors from COPFS and an Advocate Depute are working with police on this ongoing investigation.
"It is standard practice that any case regarding politicians is dealt with by prosecutors without the involvement of the Lord Advocate or Solicitor General. All Scotland's prosecutors act independently of political interference.
"As is routine, to protect the integrity of ongoing investigations, we do not comment in detail on their conduct."
A Police Scotland spokesperson said: "As this investigation is ongoing we are unable to comment further.
"The matter is active for the purposes of the Contempt of Court Act 1981 and the public are therefore advised to exercise caution if discussing it on social media."
- BBC